Smith, who broke his hand in the Test series against Pakistan in the United Arab Emirates last month, will take his place at the top of the South African order.
"I made a much quicker recovery than expected and I'm really excited to come through because this is going to be a big series to be part of," Smith said.
"We've flitted between number one and number two in the rankings over the last couple of years, so this is a really exciting opportunity for us.
"We pride ourselves on performing in these big series, there's a lot of quality on both sides and even on the field, it's exciting to be part of it," Smith added.
Teams (from):
South Africa: Graeme Smith, Alviro Petersen, Hashim Amla, Jacques Kallis, AB de Villiers, Ashwell Prince, Mark Boucher, Ryan McLaren or Lonwabo Tsotsobe, Dale Steyn, Morne Morkel, Paul Harris.
India: Gautam Gambhir, Virender Sehwag, Rahul Dravid, Sachin Tendulkar, Vangipurappu Laxman, Suresh Raina, Mahendra Singh Dhoni, Harbhajan Singh, Zaheer Khan or Jaidev Unadkat, Shanthakumaran Sreesanth, Ishant Sharma.
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